How do plants make a living? Some plants are gamblers, others are swindlers. Some plants are habitual spenders while others are strugglers and miserly savers. Plants have evolved a spectacular array of solutions to the existential problems of survival and reproduction in a world where resources are scarce, disturbances can be deadly, and competition is cut-throat. Few topics have both captured the imagination and furrowed the brows of plant ecologists, yet no topicis more important for understanding the assembly of plant communities, predicting plant responses to global change, and enhancing the restoration of our rapidly degrading biosphere. The vast array of plant strategy models that characterize the discipline now require synthesis. These models tend toemphasize either life history strategies based on demography, or functional strategies based on ecophysiology. Indeed, this disciplinary divide between demography and physiology runs deep and continues to this today. The goal of this